President Emeritus; Judge Shirley Hufstedler Professor of Biology
B.A., Swarthmore College, 1960; Ph.D., Rockefeller University, 1964; D.Sc.h.c., Swarthmore College; Mount Holyoke College; Mount Sinai Medical Center; Bard College; University of Helsinki; Cold Spring Harbor Laboratory; University of Alabama; California Polytechnic State University; Columbia University; Yale University; Rockefeller University; Harvard University; D.Phil.h.c., Weizmann Institute of Science. Professor, Caltech, 1997-2006; Millikan Professor, 2006-2021; Distinguished Professor, 2021; Hufstedler Professor, 2021-; President, 1997-2006; President Emeritus, 2006-.
